{ "abstract": "Although habitat heterogeneity is known to enhance local species diversity, the effects of management-driven structural heterogeneity on understorey plant communities across spatial scales remain poorly understood, despite their crucial role for forest biodiversity and ecosystem functioning. To analyse how forest understorey plant communities respond to an enhancement of structural heterogeneity in managed forests, we established 11 experimental sites consisting of two paired forest landscapes, an untreated homogenous control and a treatment district (ESBC). In treatment districts, structural heterogeneity was enhanced through different combinations of local patch-scale manipulations of light and deadwood features, leading to greater between-patch heterogeneity at the landscape scale. We performed a meta-analysis across these 11 sites using a Hill-Chao number and sample coverage standardisation framework. Gamma diversity increased across taxonomic, functional and phylogenetic facets in structurally heterogeneous forests (ESBC districts) via higher alpha diversity. This effect was positively associated with heterogeneity in light availability between forest patches, but not with their mean light availability. In contrast, we found no support that species turnover among patches (i.e., beta diversity) significantly contributes on average to the observed increase in gamma diversity. However, both the direction and magnitude of beta diversity responses varied substantially among landscapes. On average, structurally heterogeneous forests supported higher species richness for both open and closed forest habitat species. Synthesis and applications: Our findings highlight the benefits of enhancing structural heterogeneity for understorey plant diversity in managed forest landscapes. Specifically, management strategies that create a spatial mosaic of interventions, such as combining single-tree removal with group felling, can increase the variety of light niches among forest patches, thereby supporting the conservation of a wide range of understorey plant species, including forest specialists.", "abstract_shown": true, "affiliations": [ "Institute of Ecology, Leuphana University of Lueneburg, Lueneburg, Germany; Institute of General Ecology and Environmental Protection, TUD Dresden University of Technology, Tharandt, Germany;", "Institute of Ecology, Leuphana University of Lueneburg, Lueneburg, Germany", "Institute of General Ecology and Environmental Protection, TUD Dresden University of Technology, Tharandt, Germany", "Copernicus institute of sustainable development, Utrecht University, The Netherlands;", "Copernicus institute of sustainable development, Utrecht University, The Netherlands", "Institute of General Ecology and Environmental Protection, TUD Dresden University of Technology, Tharandt, Germany;", "Department Silviculture and Forest Ecology of the Temperate Zones, University of Goettingen, Germany;", "Department Silviculture and Forest Ecology of the Temperate Zones, University of Goettingen, Germany", "Bavarian Forest Nationalpark", "Fungal Ecology , University of Bayreuth , Universitätsstraße 30 , Bayreuth , Germany", "Fungal Ecology, University of Bayreuth, Universitaetsstrasse 30, Bayreuth, Germany", "Department of Biological Sciences, University of Toronto Scarborough, Toronto, ON, Canada;", "Department of Biological Sciences, University of Toronto Scarborough, Toronto, ON, Canada", "Institute of Statistics, National Tsing Hua University, Hsinchu, Taiwan;", "Institute of Statistics, National Tsing Hua University, Hsinchu, Taiwan", "Institute of Ecology , Leuphana University of Lüneburg , Lüneburg , Germany", "Chair of Conservation Biology and Forest Ecology, Biocenter of the Julius-Maximilians University of Wuerzburg;", "Chair of Conservation Biology and Forest Ecology, Biocenter of the Julius-Maximilians University of Wuerzburg", "Research Center for Advanced Science and Technology, The University of Tokyo, Tokyo, Japan;", "Research Center for Advanced Science and Technology, The University of Tokyo, Tokyo, Japan", "Chair of Conservation Biology and Forest Ecology , Biocenter of the Julius-Maximilians University of Würzburg", "Plant Ecology and Nature Conservation Group, Wageningen, The Netherlands", "Institute of Ecology, Leuphana University of Lueneburg, Lueneburg, Germany;" ], "article_type": null, "arxiv": null, "author_total": 16, "authors": [ { "affs": [ 0, 1, 2 ], "name": "Pia M. Bradler", "orcid": "0000-0002-9641-0519" }, { "affs": [ 3, 4 ], "name": "Benjamin M. Delory", "orcid": "0000-0002-1190-8060" }, { "affs": [ 5, 2 ], "name": "Sebastian Dittrich", "orcid": "0000-0002-4043-508X" }, { "affs": [ 6, 7 ], "name": "Christian Ammer", "orcid": "0000-0002-4235-0135" }, { "affs": [ 8, 9, 10 ], "name": "Claus Bässler", "orcid": "0000-0001-8177-8997" }, { "affs": [ 11, 12 ], "name": "Marc W. Cadotte", "orcid": "0000-0002-5816-7693" }, { "affs": [ 13, 14 ], "name": "Anne Chao", "orcid": "0000-0002-4364-8101" }, { "affs": [ 13, 14 ], "name": "Po-Yen Chuang", "orcid": "0009-0002-4993-9919" }, { "affs": [ 15, 1 ], "name": "Werner Härdtle", "orcid": "0000-0002-5599-5792" }, { "affs": [ 16, 17 ], "name": "Oliver Mitesser", "orcid": "0000-0002-3607-877X" }, { "affs": [ 18, 19 ], "name": "Akira S. Mori", "orcid": "0000-0002-8422-1198" }, { "affs": [ 8, 20, 17 ], "name": "Jörg C. Müller", "orcid": "0000-0002-1409-1586" }, { "affs": [ 18, 19 ], "name": "Keita Nishizawa", "orcid": "0000-0001-9772-0730" }, { "affs": [ 21 ], "name": "Fons van der Plas", "orcid": "0000-0003-4680-543X" }, { "affs": [ 5, 2 ], "name": "Goddert von Oheimb", "orcid": "0000-0001-7408-425X" }, { "affs": [ 22, 1 ], "name": "Andreas Fichtner", "orcid": "0000-0003-0499-4893" } ], "bibtex": "https://research.draconitic.com/paper/1rjrf09scc5kw.bib", "cited_by": 5, "doi": "10.1111/1365-2664.70477", "fulltext_url": null, "id": "1rjrf09scc5kw", "issns": [ "0021-8901", "1365-2664" ], "landing_url": "https://doi.org/10.1111/1365-2664.70477", "language": "en", "licence": "CC-BY", "licence_label": "CC BY", "licence_source": "journal", "licence_url": "https://creativecommons.org/licenses/by/4.0/", "metadata_licence": null, "native_subjects": [ [ "openalex", "2309|Ecology and Vegetation Dynamics Studies" ] ], "openalex": "W4413928349", "pdf_url": null, "pmcid": null, "pmid": null, "published": "2026-07-01", "publisher": "Wiley", "ris": "https://research.draconitic.com/paper/1rjrf09scc5kw.ris", "sources": [ { "first_seen": "2026-10-30T13:26:58Z", "native_id": "W4413928349", "source": "openalex", "url": "https://doi.org/10.1111/1365-2664.70477" } ], "subjects": [ { "field": "biology", "origin": "openalex:2309|Ecology and Vegetation Dynamics Studies", "subfield": "ecology" } ], "title": "Enhancing structural heterogeneity in managed forest landscapes promotes gamma but not beta diversity in understorey plant communities", "title_text": "Enhancing structural heterogeneity in managed forest landscapes promotes gamma but not beta diversity in understorey plant communities", "type": "peer-reviewed", "updated_at": "2026-11-08T13:20:55Z", "url": "https://research.draconitic.com/paper/1rjrf09scc5kw", "venue": "Journal of Applied Ecology", "versions": [] }